Index to
Ferns and Lycophytes of Japan

Family: Athyriaceae メシダ科 > Genus: Athyrium Roth メシダ属
GreenList Taxon ID: 517

Accepted name

Athyrium spinulosum (Maxim.) Milde

カラフトミヤマシダ

和名別名: ミヤマイヌワラビ


Basionym and homotypic synonym(s)

Cystopteris spinulosa Maxim., Mém. Pres. Acad. Imp. Sci. St.-Pétersbourg 9: 340, 1858. BHL Tropicos
Athyrium spinulosum (Maxim.) Milde, Bot. Zeitung (Berlin) 24: 376, 1866. BHL Tropicos
Asplenium spinulosum (Maxim.) Miq., Ann. Mus. Bot. Lugduno-Batavi 3(6): 174, 1867. BHL Tropicos
Pseudocystopteris spinulosa (Maxim.) Ching, Acta Phytotax. Sin. 9(1): 78, 1964. Tropicos

Type information in the protologue:
frf./frf. Maack/frf. (Am untern Amur: in feuchtem Laubwalde bei Beller, haufig/Tschjutscha/Messur, in gemischetem Walde am Abhange des Chochzier Gebriges, ?.6.30/?.8.4/1855.6.30)
Digitized type material:
Herbarium & Reg. No. Type status Collector & No. Locality Coll. date
GH00283976IsosyntypeR. Maack, #s.n.Mandshuria, ad fl. Amur1855
GH00283977IsosyntypeC. J. Maximowicz, #s.n.Amur1855
K001089294IsosyntypeC. J. Maximowicz, #s.n.Amur
U0007359IsosyntypeC. J. Maximowicz, #s.n.Amur

Information on ploidy level and reproductive mode

Chromosome number Ploidy level Reproductive mode Locality Remarks Reference
n=40, 2n=802xsexMt.Mitsutoge, Yamanashi Pref.Kurita 1976 (La Kromosomo II 2: 69-76)
n=402xsexMt.Yatsugatake, Yamanashi Pref.Hirabayashi 1970b (J. Jap. Bot. 45: 45-52)