Index to
Ferns and Lycophytes of Japan

Family: Thelypteridaceae ヒメシダ科 > Genus: Thelypteris Adans. ヒメシダ属
GreenList Taxon ID: 396

Accepted name

Thelypteris cystopteroides (D.C.Eaton) Ching

ヒメハシゴシダ

Basionym and homotypic synonym(s)

Athyrium cystopteroides D.C.Eaton, Proc. Amer. Acad. Arts 4: 110, 1858. BHL Tropicos
Asplenium cystopteroides (D.C.Eaton) Hook., Sp. Fil. 3: 220, 1860. BHL Tropicos
Dryopteris cystopteroides (D.C.Eaton) Kodama ex Tagawa, Acta Phytotax. Geobot. 3(1): 28, 1934. Link Tropicos
Thelypteris cystopteroides (D.C.Eaton) Ching, Bull. Fan Mem. Inst. Biol., Bot. 6(5): 316, 1936. Tropicos
Lastrea cystopteroides (D.C.Eaton) Copel., Gen. Fil. [Copeland] 138, 1947. Link Tropicos
Parathelypteris cystopteroides (D.C.Eaton) Ching, Acta Phytotax. Sin. 8(4): 302, 1963. Tropicos
Amauropelta cystopteroides (D.C.Eaton) S.E.Fawc. et A.R.Sm., Sida, Bot. Misc. 59: 28, 2021. Tropicos

Type information in the protologue:
(Ousima, katonasima, and Anakerima, Loo Choo islands)
Digitized type material:
Herbarium & Reg. No. Type status Collector & No. Locality Coll. date
US00134937Holotype?Wright, C., #s.n.Ousima1853
B 20 0146891IsotypeWright, C.Ousima
B 20 0146892IsotypeWright, C., #s.n.Ousima & Anakerima
GH00020629IsotypeWright, C., #s.n.Ousima, Amakerina, & Tokunosima1853--1856
K000951482IsotypeWright, C., #s.n.Ousima & [Amakerina]1853
K000951483IsotypeWright, C., #s.n.Nansei-Shoto Ousima & [Amakerima]1853
L0052272IsotypeWright, C., #s.n.Japonia
NY00128099IsotypeWright, C., #s.n.Tukuno-shima or Katonasima1853--1856

Heterotypic synonym(s) and/or incorporated taxa (forma or lower)

Athyrium hyalostegium Copel., Philipp. J. Sci. 1 (Suppl. 4): 253, 1906. BHL Tropicos
Thelypteris glanduligera (Kunze) Ching var. hyalostegia (Copel.) H.Itô, Bot. Mag. (Tokyo) 52(623): 589, 1938. Link Tropicos

Type information in the protologue:
Copeland 2033, Merrill 5186 (Luzon, ad montem Mariveles, 1350 m s.m.)
Digitized type material:
Herbarium & Reg. No. Type status Collector & No. Locality Coll. date
B 20 0028911IsosyntypeE.B. Copeland, #2033Mt. Marviles, 1350 meters1906-4
MICH1190114IsosyntypeE.B. Copeland, #2033Luzon, Bataan, Mt. Mariveles, 1350 meters1906-4
P00643475IsosyntypeE.B. Copeland, #2033Mt. Mariveles, terrestrial near summit, 1350 meters1906-4
P00643476IsosyntypeE.B. Copeland, #2033Mt. Mariveles, terrestrial near summit, 1350 meters1906-4
S-P-7555IsosyntypeE.B. Copeland, #2033Mt. Mariveles, 1350 meters1906-4
B 20 0028912Syntype Merrill, E.D., #5186 Mt. Mariveles; Prov. Bataan; Luzon1906-5
MICH1190114SyntypeE.B. Copeland, #2033Luzon, Bataan, Mt. Mariveles1906 04

Dryopteris gracilescens (Blume) Kuntze var. abbreviata Kodama, Icon. Pl. Koisik. 2(2): 43, pl. 106, 1914. Link Tropicos
Dryopteris abbreviatipinna Makino et Ogata, J. Jap. Bot. 6(7): 10, 1929. Link Tropicos

Type information in the protologue:
(Japonia australe)

Information on ploidy level and reproductive mode

Chromosome number Ploidy level Reproductive mode Locality Remarks Reference
n=ca.272xsexCultivated in Tokyo Univ. of EducationMitui 1975 (Bull. Nippon Dental Coll., General Edu. 4: 221-271)
2n=542xAmamioshima Isl., Kagoshima Pref.Kurizono 1987 (La Kromosomo II 46: 1513-1520)
n=54, 2n=1084xsexYakushima Isl., Kagoshima Pref.Kurita 1976 (La Kromosomo II 2: 69-76)
2n=542xAmami-oshima Isl.Nakato 1998a (J. Jap. Bot. 73: 314-318)
2n=542xKagoshimaNakato 2002 (Acta Phytotax. Geobot. 53: 63-76)
2n=1084xHiroshimaNakato 2002 (Acta Phytotax. Geobot. 53: 63-76)
2n=1084xMiyajima Isl., Hiroshima Pref.Kurizono 1987 (La Kromosomo II 46: 1513-1520)