Family: Lycopodiaceae ヒカゲノカズラ科 > Genus: Huperzia Bernh. コスギラン属
GreenList Taxon ID: 3
Accepted name
Huperzia serrata (Thunb.) Trevis.
トウゲシバ
和名別名: ホソバトウゲシバ
Basionym and homotypic synonym(s)
Lycopodium serratum Thunb., Syst. Veg. (ed. 14) 944, 1784. BHL Tropicos
≡ Huperzia serrata (Thunb.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248, 1875. BHL Tropicos
≡ Urostachys serratus (Thunb.) Herter, Bot. Arch. 3: 13, 1923, nom. illeg. BHL Tropicos
≡ Urostachys serratus (Thunb.) Herter, Bärlappgewächse 56, 1939, nom. illeg. Tropicos
≡ Huperzia selago (L.) Bernh. ex Schrank et Mart. subsp. serrata (Thunb.) Á.Löve et D.Löve, Univ. Colorado Stud., Ser. Biol. 17: 5, 1965. Link
≡ Huperzia selago (L.) Bernh. ex Schrank et Mart. var. serrata (Thunb.) Á.Löve et D.Löve, Univ. Colorado Stud. 17: 5, 1965. Tropicos
Type information in the protologue:
Thunb (jap.)Designated type:
Lectotype - C. P. Thunberg s.n. (JAPAN [“e Japonia”]) [UPS-THUNB 25333] designated by T. Nakaike in New Fl. Jap. Pterid. 30, 1982.Digitized type material:
| Herbarium & Reg. No. | Type status | Collector & No. | Locality | Coll. date |
|---|---|---|---|---|
| UPS-THUNB 25333B | Holotype | Thunberg | E Japonia |
Heterotypic synonym(s) and/or incorporated taxa (forma or lower)
"オニトウゲシバ"
Lycopodium serratum Thunb. var. longipetiolatum Spring, Monogr. Lycop. 2: 18, 1850 (original: 'longe-petiolatum'). BHL Tropicos
≡ Urostachys serratus (Thunb.) Herter var. longipetiolatus (Spring) Masam., Sci. Rep. Kanazawa Univ. 1: 34, 1951, nom. illeg. Link
≡ Huperzia serrata (Thunb.) Trevis. f. longipetiolata (Spring) Ching, Acta Bot. Yunnan. 3(3): 294, 1981. Tropicos
≡ Huperzia serrata (Thunb.) Trevis. var. longipetiolata (Spring) H.M.Chang, Taiwania 46(4): 380, 2001. Link Tropicos
Type information in the protologue:
([no info])Lycopodium serratum Thunb. var. thunbergii Makino, Bot. Mag. (Tokyo) 12(131): (12), 1898 (original: 'a Thunbergii'). BHL Tropicos
≡ Urostachys serratus (Thunb.) Herter var. thunbergii (Makino) Herter ex Nessel, Bärlappgewächse 59, 1939, nom. illeg.
Type information in the protologue:
((no info)) [herbarium not indicated]"チャボトウゲシバ"
Lycopodium serratum Thunb. var. alpestre H.Christ, Bull. Acad. Int. Géogr. Bot. 16(199-201): 141, 1906. BHL Tropicos
Type information in the protologue:
J. Veitch 5408a (W. CH.)Lycopodium serratum Thunb. var. integrifolium Matsuda, Bot. Mag. (Tokyo) 25(291): 124, 1911. Link
≡ Lycopodium integrifolium (Matsuda) Matsuda et Nakai ex Nakai, Bot. Mag. (Tokyo) 28(328): 102, 1914. BHL
≡ Urostachys serratus (Thunb.) Herter var. integrifolium (Matsuda) Herter ex Nessel, Bärlappgewächse 57, 1939, nom. illeg.
Type information in the protologue:
(Kiang-si: the Lu-shan)"ヒロハトウゲシバ"
Lycopodium serratum Thunb. f. intermedium Nakai, Bot. Mag. (Tokyo) 39(463): 196, 1925. Link Tropicos
≡ Lycopodium serratum Thunb. var. intermedium (Nakai) Miyabe et Kudô, J. Fac. Agric. Hokkaido Imp. Univ. 26(1): 53, 1930. Link
≡ Huperzia serrata (Thunb.) Trevis. f. intermedia (Nakai) Ching, Acta Bot. Yunnan. 3(3): 294, 1981. Tropicos
≡ Huperzia serrata (Thunb.) Trevis. var. intermedia (Nakai) Satou, Hikobia 12(3): 268, 1997.
Type information in the protologue:
C. J. Maximowicz/U. Faurie 3904/U. Faurie/Savatier 1525 (Yeso: Fukuyama/Yeso: Fukuyama/Hondo: Aomori/Hondo: Yokosuka) [herbarium not indicated (syn)]Urostachys serratus (Thunb.) Herter var. kiyosumiyamaensis Onuma ex Nessel, Bärlappgewächse 60, 1939, nom. illeg.
Type information in the protologue:
()"トヨグチトウゲシバ"
Lycopodium serratum Thunb. f. paucidentatum Sugim., Keys Herb. Pl. Jap. Pterid. 88, 409, 1966.
≡ Huperzia serrata (Thunb.) Trevisan var. paucidentata (Sugim.) Satou, Hikkobia 12(3): 268, 1997.
Type information in the protologue:
J. Sugimoto (Hondo: Prov. Shinano, Mt. Toyoguchi, 1955) [Hb. Sugimoto, Shizuoka (holo)]Information on ploidy level and reproductive mode
| Chromosome number | Ploidy level | Reproductive mode | Locality | Remarks | Reference |
|---|---|---|---|---|---|
| n=68 | 4x | Uchiurayama, Chiba Pref. | as var. intermedium (Nakai) Miyabe et Kudô | Takamiya & Kurita 1983 (Acta Phytotax. Geobot. 34: 66-79) | |
| n=136 | 8x | Uchiurayama, Chiba Pref.; Mt.Ogasayama, Shizuoka Pref.; Kudosan-cho, Wakayama Pref.; Yakushima Isl., Kagoshima Pref. | as var. intermedium (Nakai) Miyabe et Kudô | Takamiya & Kurita 1983 (Acta Phytotax. Geobot. 34: 66-79) | |
| n=68 | 4x | Mt.Tanigawadake, Gunma Pref.; Mt.Kurohimeyama, Nagano Pref.; Mt.Ishizuchiyama, Ehime Pref. | as var. serratum | Takamiya & Kurita 1983 (Acta Phytotax. Geobot. 34: 66-79) | |
| 2n=204 | 6x | Mt.Shiratoriyama, Miyazaki Pref. | as var. serratum | Takamiya 1984 (Chromosome Infor. Serv. 35: 27-28) | |
| n=136 | 8x | Uchiurayama & Sekisonsan, Chiba Pref.; Yugashima-cho & Mt.Ogasayama, Shizuoka Pref. | as var. serratum | Takamiya & Kurita 1983 (Acta Phytotax. Geobot. 34: 66-79) |