Index to
Ferns and Lycophytes of Japan

Family: Lycopodiaceae ヒカゲノカズラ科 > Genus: Huperzia Bernh. コスギラン属
GreenList Taxon ID: 3

Accepted name

Huperzia serrata (Thunb.) Trevis.

トウゲシバ

和名別名: ホソバトウゲシバ


Basionym and homotypic synonym(s)

Lycopodium serratum Thunb., Syst. Veg. (ed. 14) 944, 1784. BHL Tropicos
Huperzia serrata (Thunb.) Trevis., Atti Soc. Ital. Sci. Nat. 17: 248, 1875. BHL Tropicos
Urostachys serratus (Thunb.) Herter, Bot. Arch. 3: 13, 1923, nom. illeg. BHL Tropicos
Urostachys serratus (Thunb.) Herter, Bärlappgewächse 56, 1939, nom. illeg. Tropicos
Huperzia selago (L.) Bernh. ex Schrank et Mart. subsp. serrata (Thunb.) Á.Löve et D.Löve, Univ. Colorado Stud., Ser. Biol. 17: 5, 1965. Link
Huperzia selago (L.) Bernh. ex Schrank et Mart. var. serrata (Thunb.) Á.Löve et D.Löve, Univ. Colorado Stud. 17: 5, 1965. Tropicos

Type information in the protologue:
Thunb (jap.)
Designated type:
Lectotype - C. P. Thunberg s.n. (JAPAN [“e Japonia”]) [UPS-THUNB 25333] designated by T. Nakaike in New Fl. Jap. Pterid. 30, 1982.
Digitized type material:
Herbarium & Reg. No. Type status Collector & No. Locality Coll. date
UPS-THUNB 25333BHolotypeThunbergE Japonia

Heterotypic synonym(s) and/or incorporated taxa (forma or lower)

"オニトウゲシバ"
Lycopodium serratum Thunb. var. longipetiolatum Spring, Monogr. Lycop. 2: 18, 1850 (original: 'longe-petiolatum'). BHL Tropicos
Urostachys serratus (Thunb.) Herter var. longipetiolatus (Spring) Masam., Sci. Rep. Kanazawa Univ. 1: 34, 1951, nom. illeg. Link
Huperzia serrata (Thunb.) Trevis. f. longipetiolata (Spring) Ching, Acta Bot. Yunnan. 3(3): 294, 1981. Tropicos
Huperzia serrata (Thunb.) Trevis. var. longipetiolata (Spring) H.M.Chang, Taiwania 46(4): 380, 2001. Link Tropicos

Type information in the protologue:
([no info])

Lycopodium serratum Thunb. var. thunbergii Makino, Bot. Mag. (Tokyo) 12(131): (12), 1898 (original: 'a Thunbergii'). BHL Tropicos
Urostachys serratus (Thunb.) Herter var. thunbergii (Makino) Herter ex Nessel, Bärlappgewächse 59, 1939, nom. illeg.

Type information in the protologue:
((no info)) [herbarium not indicated]

"チャボトウゲシバ"
Lycopodium serratum Thunb. var. alpestre H.Christ, Bull. Acad. Int. Géogr. Bot. 16(199-201): 141, 1906. BHL Tropicos

Type information in the protologue:
J. Veitch 5408a (W. CH.)

Lycopodium serratum Thunb. var. integrifolium Matsuda, Bot. Mag. (Tokyo) 25(291): 124, 1911. Link
Lycopodium integrifolium (Matsuda) Matsuda et Nakai ex Nakai, Bot. Mag. (Tokyo) 28(328): 102, 1914. BHL
Urostachys serratus (Thunb.) Herter var. integrifolium (Matsuda) Herter ex Nessel, Bärlappgewächse 57, 1939, nom. illeg.

Type information in the protologue:
(Kiang-si: the Lu-shan)

"ヒロハトウゲシバ"
Lycopodium serratum Thunb. f. intermedium Nakai, Bot. Mag. (Tokyo) 39(463): 196, 1925. Link Tropicos
Lycopodium serratum Thunb. var. intermedium (Nakai) Miyabe et Kudô, J. Fac. Agric. Hokkaido Imp. Univ. 26(1): 53, 1930. Link
Huperzia serrata (Thunb.) Trevis. f. intermedia (Nakai) Ching, Acta Bot. Yunnan. 3(3): 294, 1981. Tropicos
Huperzia serrata (Thunb.) Trevis. var. intermedia (Nakai) Satou, Hikobia 12(3): 268, 1997.

Type information in the protologue:
C. J. Maximowicz/U. Faurie 3904/U. Faurie/Savatier 1525 (Yeso: Fukuyama/Yeso: Fukuyama/Hondo: Aomori/Hondo: Yokosuka) [herbarium not indicated (syn)]

Urostachys serratus (Thunb.) Herter var. kiyosumiyamaensis Onuma ex Nessel, Bärlappgewächse 60, 1939, nom. illeg.

Type information in the protologue:
()

Lycopodium serratum Thunb. f. paucidentatum Sugim., Keys Herb. Pl. Jap. Pterid. 88, 409, 1966.
Huperzia serrata (Thunb.) Trevisan var. paucidentata (Sugim.) Satou, Hikkobia 12(3): 268, 1997.

Type information in the protologue:
J. Sugimoto (Hondo: Prov. Shinano, Mt. Toyoguchi, 1955) [Hb. Sugimoto, Shizuoka (holo)]

Information on ploidy level and reproductive mode

Chromosome number Ploidy level Reproductive mode Locality Remarks Reference
n=684xUchiurayama, Chiba Pref.as var. intermedium (Nakai) Miyabe et KudôTakamiya & Kurita 1983 (Acta Phytotax. Geobot. 34: 66-79)
n=1368xUchiurayama, Chiba Pref.; Mt.Ogasayama, Shizuoka Pref.; Kudosan-cho, Wakayama Pref.; Yakushima Isl., Kagoshima Pref.as var. intermedium (Nakai) Miyabe et KudôTakamiya & Kurita 1983 (Acta Phytotax. Geobot. 34: 66-79)
n=684xMt.Tanigawadake, Gunma Pref.; Mt.Kurohimeyama, Nagano Pref.; Mt.Ishizuchiyama, Ehime Pref.as var. serratumTakamiya & Kurita 1983 (Acta Phytotax. Geobot. 34: 66-79)
2n=2046xMt.Shiratoriyama, Miyazaki Pref.as var. serratumTakamiya 1984 (Chromosome Infor. Serv. 35: 27-28)
n=1368xUchiurayama & Sekisonsan, Chiba Pref.; Yugashima-cho & Mt.Ogasayama, Shizuoka Pref.as var. serratumTakamiya & Kurita 1983 (Acta Phytotax. Geobot. 34: 66-79)