Index to
Ferns and Lycophytes of Japan

Family: Polypodiaceae ウラボシ科 > Genus: Lepisorus (J.Sm.) Ching ノキシノブ属
GreenList Taxon ID: 1054

Accepted name

Lepisorus thunbergianus (Kaulf.) Ching

ノキシノブ

Basionym and homotypic synonym(s)

Polypodium lineare Thunb., Syst. Veg. (ed. 14) 934, 1784, nom. illeg. BHL Tropicos
Pleopeltis thunbergiana Kaulf., Wesen Farrenk. 113, 1827. Link Tropicos
Pleopeltis linearis (Thunb.) T.Moore, Index Fil. [Moore] 346, 1862. BHL Tropicos
Polypodium lineare Thunb. var. thunbergianum (Kaulf.) Takeda, Notes Roy. Bot. Gard. Edinburgh 8(39): 268, 1915. BHL Tropicos
Lepisorus thunbergianus (Kaulf.) Ching, Bull. Fan Mem. Inst. Biol. 4(3): 88, 1933. Tropicos
Polypodium thunbergianum (Kaulf.) C.Chr., Index Filic., Suppl. 3 160, 1934. BHL Tropicos

Type information in the protologue:
Thunb. (Kosido)
Designated type:
Lectotype - C. P. Thunberg s.n. (JAPAN [“e Japonia”]) [UPS-THUNB 24513] designated by T. Nakaike in New Fl. Jap. Pterid. 668, 1982.
Digitized type material:
Herbarium & Reg. No. Type status Collector & No. Locality Coll. date
UPS-THUNB 24513Holotypeunknown (C.P. Thunberg?)Japan

Heterotypic synonym(s) and/or incorporated taxa (forma or lower)

Polypodium lineare Thunb. f. caudatoattenuatum Takeda, Notes Roy. Bot. Gard. Edinburgh 8(39): 269, 1915 (original: 'caudato-attenuatum'). BHL Tropicos
Polypodium caudatoattenuatum (Takeda) C.Chr., Index Filic., Suppl. 3 146, 1934. BHL Tropicos

Type information in the protologue:
Savatier 1541 part/C. Wright, 1853-1856, ex hb. U.S. Pacif. Expl. Exped. (Japan. Yokosuka/Japan: Oshima)

Information on ploidy level and reproductive mode

Chromosome number Ploidy level Reproductive mode Locality Remarks Reference
n=25, 2n=502xsexKiyosumiyama, Chiba Pref.Takei 1982a (Res. Bull. Fac. Educ. Oita Univ. (Nat. Sci.) 6(2): 1-21)
n=252xsexHashimoto, Kanagawa Pref.Mitui 1966a (J. Jap. Bot. 41: 60-64)
n=252xsexHashimoto, Enoshima, Kamakura City, & Oyama, Kanagawa Pref.Mitui 1966b (J. Jap. Bot. 41: 270-276)
n=252xsexTaura, Kanagawa Pref.Mitui 1968 (Sci. Rep. Tokyo Kyoiku Daigaku, Sec. B 13: 285-333)
n=252xsexHigashisakurajima, Kagoshima Pref.Mitui 1975 (Bull. Nippon Dental Coll., General Edu. 4: 221-271)
2n=502xsexOshima Isl., Tokyo Pref.Mitui 1971 (J. Jap. Bot. 46: 83-96)
2n=502xTottori City & Iwami-machi, Tottori Pref.; Shiroyama, Ichihatayakushi & Taisya-machi, Shimane Pref.; Onomichi City, Sakamachi, Kurahashi-machi, Mukoujina Isl., Itsukaichi-machi, Futsukaichi-machi, Ono-machi, & Miyajima Isl., Hiroshima Pref.; Omijima Isl., Yamaguchi Pref.; Mitsuhama & Shiroyama, Ehime Pref.; Yoshiragawa, Kochi Pref.; Nichinankaigan, Miyazaki Pref.; Mt.Kaimondake & Yakushima Isl., Kagoshima Pref.Takei 1974 (J. Jap. Bot. 49: 356-359)
2n=502xsexOshima Isl., Tokyo Pref.Mitui 1975 (Bull. Nippon Dental Coll., General Edu. 4: 221-271)
2n=502xWakari, Kanda-machi, & Yukuhashi City, Fukuoka Pref.; Higashinakatsu, Usa City, Takada City, Tateishi, Aki, Kitsuki City, Honyabakei, Oda, Amagase-machi, Hita City, Yufuin-machi, Minamiyufu, Yudaira, Nukaigahara, Kasuga-machi, Dannobara, Reizan, Okajoshi, Ogi-machi, Inukai-machi, & Mie-machi, Oita Pref.Takei 1978 (J. Jap. Bot. 53: 87-91)
2n=502xHiroshima City, Hiroshima Pref.; Matsuyama City, Ehime Pref.; Dannoharu & Takeda City, Oita Pref.Takei 1982b (Res. Bull. Fac. Educ. Oita Univ. (Nat. Sci.) 6(3): 9-41)
2n=502xOkutama, Tokyo Pref.; Oyama & Kamakura City, Kanagawa Pref.; Kashiwa City, Mitsuishiyama, Minato, Ohara, Katsuura, & Kiyosumiyama, Chiba Pref.Nakato et al. 1983 (J. Jap. Bot. 58: 195-205)
2n=502xMiyakejima Isl. & Hachijojima Isl., Tokyo Pref.; Ito, Yugashima, & Heda, Shizuoka Pref.; Kaifu, Tokushima Pref.; Kumamoto City, Kumamoto Pref.; Yakushima Isl., Kagoshima Pref.; Iriomotejima Isl., Okinawa Pref.Mitui et al. 1987 (J. Jap. Bot. 62: 311-319)
2n=502xsexOyama, Kanagawa Pref.; Yugashima, Shizuoka Pref.Masuyama et al. 1987 (J. Jap. Bot. 62: 321-331)
2n=502xOita City, Oita Pref.Takei 1988c (Res. Bull. Fac. Educ. Oita Univ. 10: 89-104)
2n=502xNo locality givenTakei 1989 (Cytologia 54: 135-144)
2n=502xNo locality givenMitui et al. 1989b (Plant Chromosome Research 1987 )
2n=512xOyama, Kanagawa Pref.Nakato et al. 1983 (J. Jap. Bot. 58: 195-205)
2n=512xNo locality givenMitui et al. 1989b (Plant Chromosome Research 1987 )
2n=753xItsukaichi-machi, Hiroshima Pref.Takei 1974 (J. Jap. Bot. 49: 356-359)
2n=753xOkutama, Tokyo Pref.Nakato et al. 1983 (J. Jap. Bot. 58: 195-205)
2n=753xNo locality givenMitui et al. 1989b (Plant Chromosome Research 1987 )
(2n=75)3xirrKawai, Tokyo Pref.Mitui 1968 (Sci. Rep. Tokyo Kyoiku Daigaku, Sec. B 13: 285-333)
2n=763xKamakura City, Kanagawa Pref.; Mitsuishiyama & Minato, Chiba Pref.Nakato et al. 1983 (J. Jap. Bot. 58: 195-205)
2n=763xNo locality givenMitui et al. 1989b (Plant Chromosome Research 1987 )
n=504xsexHanno City, Saitama Pref.Mitui 1965 (J. Jap. Bot. 40: 117-124)
n=514xsexKawamata, Saitama Pref.; Otaba, Tokyo Pref.Mitui 1966b (J. Jap. Bot. 41: 270-276)
n=524xsexKawai, Tokyo Pref.Mitui 1968 (Sci. Rep. Tokyo Kyoiku Daigaku, Sec. B 13: 285-333)
n=534xsexShinshiro, Aichi Pref.Mitui 1971 (J. Jap. Bot. 46: 83-96)
n=544xsexShibata, Niigata Pref.Mitui 1975 (Bull. Nippon Dental Coll., General Edu. 4: 221-271)
2n=1004xIkimaeshima Isl. & Yamaguchi-machi, Shimane Pref.; Fuchu-cho, Asa-machi, Koi-machi, Yamada-machi, Yoshiwa-mura, Yuki-machi, & Takei 1974 (J. Jap. Bot. 49: 356-359)
2n=1004xYabakei-machi, Sikakura, Oda, Amagase-machi, Yufuin-machi, Minamiyufu, & Yunodaira, Oita Pref.Takei 1978 (J. Jap. Bot. 53: 87-91)
2n=1004xAshiyasu-mura, Yamanashi Pref.; Yasufuruichi, Hiroshima Pref.; Yabakei, Oita Pref.Takei 1982b (Res. Bull. Fac. Educ. Oita Univ. (Nat. Sci.) 6(3): 9-41)
2n=1004xTomioka City, Gunma Pref.; Yorii, Saitama Pref.; Okutama, Tokyo Pref.; Oyama & Kamakura City, Kanagawa Pref.; Kashiwa City & Minato, Chiba Pref.Nakato et al. 1983 (J. Jap. Bot. 58: 195-205)
2n=1004xAkita City, Akita Pref.; Ohtaba & Itsukaichi, Tokyo Pref.; Mt.Daisen, Tottori Pref.Mitui et al. 1987 (J. Jap. Bot. 62: 311-319)
2n=1004xNo locality givenMitui et al. 1989b (Plant Chromosome Research 1987 )
2n=1014xOkutama, Tokyo Pref.; Oyama, Kanagawa Pref.Nakato et al. 1983 (J. Jap. Bot. 58: 195-205)
2n=1014xIkawa, Akita Pref.Mitui et al. 1987 (J. Jap. Bot. 62: 311-319)
2n=1014xNo locality givenMitui et al. 1989b (Plant Chromosome Research 1987 )
2n=1024xAsa-machi & Yuki-machi, Hiroshima Pref.Takei 1974 (J. Jap. Bot. 49: 356-359)
2n=1024xShinyabakei, Hosenji, & Uchiyama, Oita Pref.Takei 1978 (J. Jap. Bot. 53: 87-91)
2n=1024xHosenji, Oita Pref.Takei 1982b (Res. Bull. Fac. Educ. Oita Univ. (Nat. Sci.) 6(3): 9-41)
2n=1024xYorii, Saitama Pref.; Okutama, Tokyo Pref.; Oyama, Kanagawa Pref.; Minato, Chiba Pref.Nakato et al. 1983 (J. Jap. Bot. 58: 195-205)
2n=1024xJuniko, Aomori Pref.; Ikawa & Akita City, Akita Pref.; Yahiko & Wajima, Niigata Pref.Mitui et al. 1987 (J. Jap. Bot. 62: 311-319)
2n=1024xNo locality givenMitui et al. 1989b (Plant Chromosome Research 1987 )
2n=1034xIkawa, Akita Pref.Mitui et al. 1987 (J. Jap. Bot. 62: 311-319)
2n=1034xNo locality givenMitui et al. 1989b (Plant Chromosome Research 1987 )
2n=502xHigashiyamato, Ome & Hinode, Tokyo Pref.Shinohara et al. 2010 (Syst. Bot. 35: 20-29)
2n=753xOme & Okutama, Tokyo Pref.Shinohara et al. 2010 (Syst. Bot. 35: 20-29)
2n=763xOme & Akiruno, Tokyo Pref.Shinohara et al. 2010 (Syst. Bot. 35: 20-29)
2n=1004xOme, Okutama, Akiruno, Hinohara & Chiyoda-ku Tokyo Pref.; Kosuge, Yamanashi Pref.Shinohara et al. 2010 (Syst. Bot. 35: 20-29)
2n=1014xOme & Okutama, Tokyo Pref.Shinohara et al. 2010 (Syst. Bot. 35: 20-29)
2n=1024xOkutama, Akiruno, Ome & Hinohara, Tokyo Pref.; Kosuge, Yamanashi Pref.Shinohara et al. 2010 (Syst. Bot. 35: 20-29)